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2037, 2041, 2061, 2079, 2084, 2113, 2124, 2136, 2144, 2174, 2182, 2185, 2190, 2193, 2199, 2202, 2210, 2214, 2219, 2224, 2237, 2257, 2277, 2289, 2341, 2346, 2352, 2357, 2372, 2378, 2386, 2395, 2398, 2406, 2409, 2415, 2423, 2430, 2434, 2441, 2456, 2463, 2466, 2469, 2473, 2497, 2500, 2507, 2511, 2518, 2521, 2541, 2576, 2587, 2600, 2616, 2636, 2653, 2660, 2666, 2692, 2734, 2739, 2742, 2751, 2760, 2772, 2775, 2786, 2791, 2809, 2833, 2865, 2880, 2889, 2893, 2901, 2916, 2920, 2927, 2950, 2955, 2971, 2996, 3000, 3011, 3033, 3037, 3042, 3052, 3055, 3072, 3088, 3094, 3116, 3124, 3128, 3132, 3135, 3154, 3158, 3163, 3167, 3176, 3237, 3252, 3256, 3262, 3279, 3284, 3290, 3293, 3344, 3347, 3350, 3354, 3365, 3370, 3416, 3428, 3438, 3443, 3453, 3464, 3467, 3475, 3497, 3515, 3526, 3533, 3537, 3545, 3560, 3563, 3573, 3583, 3597, 3602, 3609, 3677, 3688, 3717, 3726, 3738, 3742, 3746, 3772, 3787, 3797, 3812, 3821, 3830, 3837, 3843, 3852, 3857, 3883, 3887, 3911, 3926, 3934, 3938, 3948, 3953, 3964, 3967, 3972, 4019, 4033, 4047, 4065, 4074, 4078, 4123, 4127, 4141, 4148, 4151, 4171, 4176, 4200, 4209, 4246, 4251, 4260, 4317, 4366, 4370, 4385, 4399, 4418, 4426, 4436, 4444, 4467, 4479, 4482, 4492, 4496, 4511], 'Wiley': [151], 'Online': [152], 'Library': [153], 'UseShareable': [158], 'LinkUse': [159], 'link': [161], 'a': [165, 180, 193, 324, 570, 600, 749, 788, 830, 894, 942, 1099, 1167, 1238, 1279, 1300, 1346, 1350, 1391, 1596, 1685, 1758, 1948, 1967, 2008, 2015, 2017, 2049, 2076, 2103, 2220, 2318, 2367, 2369, 2452, 2481, 2489, 2566, 2626, 2695, 2699, 2720, 2737, 2767, 2800, 2822, 2959, 2985, 3016, 3277, 3305, 3326, 3387, 3447, 3569, 3683, 3712, 3720, 3734, 3751, 3804, 3905, 4205, 4326, 4362, 4396, 4414, 4432, 4440, 4454, 4463, 4504], 'article': [170, 969, 1365, 2691], 'with': [171, 297, 339, 538, 913, 1649, 1772, 2116, 2702, 4071, 4080, 4215, 4328, 4373, 4435], 'your': [172], 'friends': [173], 'colleagues.': [175], 'Learn': [176], 'more.Copy': [177], 'URL': [178], 'Share': [179], 'linkShare': [181], 'onFacebookTwitterLinked': [182], 'InRedditWechat': [183], 'Abstract': [184], 'The': [185, 346, 740, 815, 965, 1171, 1364, 1502, 1508, 1578, 1866, 1962, 1999, 2247, 2260, 2418, 2852, 3028, 3220, 3248, 3660, 3943, 4221, 4293], 'goal': [186], 'study': [189], 'is': [190, 314, 369, 477, 520, 553, 595, 658, 716, 724, 779, 869, 919, 924, 970, 1013, 1088, 1160, 1176, 1366, 1417, 1420, 1514, 1527, 1533, 1554, 1568, 1592, 1655, 1722, 1746, 1751, 1757, 1785, 1812, 1817, 1880, 1890, 1966, 2023, 2050, 2119, 2196, 2254, 2329, 2389, 2401, 2426, 2449, 2478, 2503, 2575, 2593, 2606, 2726, 2758, 2868, 2943, 3015, 3045, 3098, 3104, 3184, 3304, 3334, 3393, 3412, 3435, 3456, 3484, 3504, 3558, 3695, 3863, 3925, 4089, 4132, 4144, 4204, 4325, 4378, 4384, 4395, 4431, 4485, 4491, 4502, 4507, 4522], 'develop': [192], 'unifying': [194, 490], 'theoretical': [195, 926, 1343], 'framework': [196], 'quantify': [198, 854, 879, 1007], 'strength': [200, 329, 979, 1009, 1017, 1051, 4400], 'reproductive': [202, 366, 587, 686, 981, 1011, 2578, 2588, 2602, 2973, 3477, 3517, 3928], 'isolation.': [203, 331, 367, 687, 982, 1107], 'We': [204, 263, 574, 983, 1039, 1370, 1437, 1471, 1635, 1714, 1845, 1900, 2091, 2404, 2683, 2803, 2904, 2947, 3100, 3259, 3310, 3385, 3511], 'propose': [205, 972], 'use': [207, 266, 535, 821, 1403, 2229, 2673], '"effective': [210, 614, 1045, 1155], 'recombination': [211, 242, 258, 309, 1024, 1046, 1081, 1156, 1218, 1243, 1310, 1357, 1378, 1398, 1492, 1627, 1742, 1783, 1851, 1868, 1885, 1911, 1916, 1920, 1926, 1964, 2126, 2187, 2239, 2668, 2952, 3002, 3013, 3137, 3160, 3193, 3440, 3539, 3575, 3626, 3722, 3773, 3862, 3874, 3913, 4008, 4058, 4178, 4202, 4270, 4493], 'rate,"': [212, 1047], 'which': [213, 425, 486, 761, 1048, 1059, 1150, 1312, 1756, 2268, 2757, 2794, 3240, 3390, 3866, 3962], 'measures': [214, 1049, 1801, 1870, 3963], 'how': [215, 427, 593, 1330, 1440, 1643], 'fast': [216], 'associations': [217, 994, 1036], 'genes': [219, 1022, 2373, 3932], 'are': [220, 444, 795, 845, 989, 1028, 1285, 1296, 1313, 1336, 1537, 1843, 1896, 2055, 2160, 2319, 2349, 2548, 2797, 2875, 2892, 2912, 3066, 3243, 3251, 3264, 3271, 3297, 3422, 3614, 3655, 3790, 3805, 3868, 4022, 4118, 4225, 4422, 4514], 'broken': [221], 'interlocus': [223, 1019], 'recombination.': [224, 3849, 4343], 'Applying': [225], 'well-established': [227], 'theory': [228, 1383, 1461, 1509, 1567, 1591], 'effective': [231, 241, 308, 654, 704, 721, 744, 875, 1080, 1085, 1113, 1217, 1242, 1309, 1356, 1361, 1377, 1386, 1397, 1491, 1626, 1850, 1867, 1910, 1919, 1963, 1975, 2000, 2033, 2062, 2125, 2186, 2203, 2238, 2342, 2353, 2522, 2533, 2560, 2667, 2951, 3001, 3012, 3136, 3159, 3192, 3439, 3538, 3574, 3625, 3912, 4177, 4201, 4269], 'migration': [232, 250, 615, 655, 660, 705, 711, 722, 730, 745, 876, 1086, 1114, 1362, 1387, 1976, 2001, 2034, 2063, 2204, 2302, 2343, 2354, 2523, 2534, 2546, 2561, 2925, 3172, 3201, 3417, 3634, 3691, 3881], 'rate,': [233, 1311, 1388, 1977], 'we': [234, 564, 998, 1174, 1284, 1348, 1389, 1428, 1455, 1600, 1623, 1825, 1979, 2082, 2141, 2228, 2269, 2624, 2647, 2672, 2687, 2789, 2886, 3058, 3122, 3169, 3235, 3680, 3699, 3710, 3779, 4249, 4311, 4360], 'derive': [235, 1390, 2233, 3711], 'two': [236, 269, 272, 1066, 1408, 1432, 1538, 1546, 1582, 1608, 1645, 1669, 1688, 1737, 1808, 1876, 1894, 2117, 2226, 2264, 2278, 2347, 2358, 2379, 2424, 2678, 2703, 2776, 3043, 3144, 3246, 3267, 3693, 3813, 4423, 4497], 'techniques': [237, 844, 1444], 'investigate': [239, 1249], 'rate': [243, 310, 656, 661, 667, 706, 712, 723, 746, 877, 1057, 1082, 1087, 1115, 1219, 1244, 1358, 1399, 1493, 1628, 1784, 1792, 1852, 1869, 1886, 1912, 1921, 1927, 1965, 2002, 2035, 2064, 2127, 2188, 2421, 2524, 2953, 3003, 3014, 3161, 3441, 3540, 3723, 3753, 3774, 4179, 4203, 4494], 'in': [244, 320, 365, 382, 447, 496, 664, 674, 736, 893, 935, 1260, 1316, 1404, 1423, 1511, 1664, 1982, 2040, 2167, 2173, 2206, 2251, 2266, 2292, 2322, 2440, 2582, 2595, 2615, 2644, 2656, 2689, 2698, 2747, 2750, 2762, 2793, 2839, 2855, 2909, 3047, 3068, 3078, 3110, 3188, 3239, 3255, 3343, 3389, 3397, 3406, 3425, 3472, 3481, 3522, 3601, 3616, 3621, 3706, 3741, 3769, 3791, 3796, 3842, 3856, 3869, 3876, 3937, 4062, 4105, 4120, 4147, 4227, 4259, 4265, 4314, 4354, 4462], 'models': [245, 443, 769, 835, 1335, 1451, 2679], 'speciation:': [247], 'weak': [249, 257, 781], 'approximation': [251, 259, 843, 1393, 1415, 2234, 2248, 2654, 2663, 2997, 3056, 3096, 3534, 3685, 3714, 3759, 3908], 'parapatric': [253, 607, 885, 907, 1332, 1405, 2681], 'scenarios': [254, 338, 1497], 'sympatric': [261, 895, 949, 1334, 1424], 'scenarios.': [262], 'illustrate': [264, 1429], 'these': [268, 1027], 'methods': [270, 794, 824, 1305], 'examples': [273, 1409, 1433, 1953, 3108], 'each:': [274], '(1)': [275, 1297, 1523, 1553, 1755, 1771, 2841], 'single-locus': [276, 3197], 'genetic': [277, 282, 891, 1106, 1197, 1531, 2375, 2383, 2714, 2795, 2862, 2933, 3241, 3331, 3410, 3756, 3809], 'incompatibility': [278, 283, 3198, 3631], '(2)': [280, 1320, 1535, 1580, 1777, 1779, 1820, 2844], 'two-locus': [281, 3630, 4345], 'first': [286, 1371, 1692, 2555, 2787, 2989, 3491, 4098], 'method,': [287], '(3)': [289, 1542, 1618, 1865, 2169, 2847, 3142], 'assortative': [290, 294, 4052, 4095, 4212, 4274, 4322], 'mating': [291, 295, 1745, 1811, 1889, 2849, 4096, 4159, 4213], '(4)': [293, 2026], 'combined': [296], 'disruptive': [298, 2392, 4334, 4376, 4402], 'selection': [299, 1210, 1750, 1816, 2881, 3429, 4377], 'second': [302, 1414, 1701, 3906], 'method.': [303], 'An': [304, 1076, 1918], 'advantage': [305, 848, 1077, 4434], 'over': [311, 1083], 'previous': [312, 1463, 3257, 3603, 3678, 3743, 4261], 'approaches': [313, 2777], 'that': [315, 376, 500, 591, 662, 873, 930, 987, 1089, 1135, 1419, 1530, 1637, 1653, 1781, 2140, 2152, 2314, 2340, 2433, 2569, 2599, 2609, 2650, 2685, 2713, 2805, 2888, 2907, 2987, 3010, 3093, 3102, 3181, 3261, 3270, 3312, 3360, 3503, 3559, 3690, 3781, 3851, 3893, 3917, 3998, 4027, 4046, 4055, 4140, 4150, 4168, 4199], 'it': [316, 519, 552, 1090, 1159, 1484, 1862, 2502, 3765, 3891, 4025, 4166], 'integrates': [317, 1092], 'gene': [318, 596, 689, 737, 880, 1035, 1093, 1124, 2100, 2508], 'flow': [319, 597, 690, 715, 738, 881, 1094, 1125, 2101, 2467, 2509, 2693], 'both': [321, 1096, 2207, 2872, 3463, 4236], 'directions': [322, 1097, 2208], 'into': [323, 1098, 1607, 2716, 2995, 3532, 4032, 4189], 'single': [325, 1100, 1168, 1342, 2801], 'index': [326, 1101, 2097, 4128], 'measuring': [327, 1102, 4398], 'This': [332, 542, 911, 2131, 2337, 3091, 3333, 3409, 3488, 3731, 3896, 4091, 4336, 4344, 4404, 4459], 'enables': [333], 'straightforward': [334], 'comparisons': [335, 544, 864], 'speciation': [337, 510, 528, 608, 834, 950, 1317, 1406, 1425], 'same': [341, 666, 2038, 2894, 3253, 3598, 4075, 4252], 'or': [342, 360, 549, 685, 782, 991, 1144, 1559, 1672, 1815, 2488, 3376], 'different': [343, 546, 866, 890, 1141, 1450, 2080, 3755], 'geographic': [344, 917, 2382, 3704], 'histories.': [345], 'method': [347], 'also': [348, 862, 888, 1127, 1263, 1438, 1594, 2093, 3508, 4162], 'allows': [349, 3898], 'us': [350, 3899, 4044], 'evaluate': [352], 'relative': [354, 1789, 4386], 'contributions': [355], 'F2': [357], 'hybrid': [358, 3565], 'deficiency': [359], 'linkage': [361, 1063, 1192, 1733, 1795, 2147, 4464, 4476, 4505], 'between': [362, 545, 865, 884, 889, 1065, 1354, 1735, 1806, 1853, 1875, 2128, 2189, 2213, 2345, 2356, 2377, 2472, 2638, 2954, 3143, 3442, 3692, 3725, 3754, 3811, 3886, 4466, 4478, 4495], 'multiple': [363], 'barriers': [364], 'Speciation': [368, 442, 1945], 'one': [370, 1071, 1615, 2515, 2607, 3282, 3319, 3739], 'fundamental': [373, 1351], 'evolutionary': [374, 433], 'puzzles': [375], 'Darwin': [377], 'essentially': [378], 'failed': [379], 'address': [381], 'Origin': [384], 'Species': [386], 'despite': [387], 'title': [389], 'book.': [392], 'Not': [393], 'mention': [395], 'empirical': [396, 928], 'researchers': [397], '(Coyne': [398], 'Orr': [400], '2004;': [401, 437, 810], 'Widmer': [402], 'et': [403, 413, 439, 469, 642, 758, 812, 957, 962, 1224, 1230, 2070, 2088], 'al.': [404, 414, 440, 470, 643, 759, 813, 958, 963, 1225, 1231, 2071, 2089], '2009),': [405], 'challenging': [407], 'problem': [408, 495, 756], 'intrigued': [409], 'many': [410, 501, 1271], 'theoreticians': [411], '(Turelli': [412], '2001;': [415], 'Gavrilets': [416, 631, 637, 3383, 3509, 4163, 4357, 4410], '2004).': [417, 441, 2074, 3384, 4164], 'They': [418], 'contributed': [419], 'numerous': [420], 'mathematical': [421, 750, 1281, 1352, 3030], 'simulation': [423], 'models,': [424], 'revealed': [426], 'species': [428, 988], 'can': [429, 565, 692, 878, 1445, 1619, 1826, 2132, 2142, 2977, 3512, 4180, 4460], 'diverge': [430], 'through': [431, 2565], 'various': [432, 1331], 'mechanisms': [434], '(Gavrilets': [435], '2003,': [436], 'Dieckmann': [438], 'still': [445, 1569, 3762], 'increasing': [446], 'number,': [448], 'continuously': [449], 'bringing': [450], 'novel': [451], 'ideas': [452], 'renewing': [454], 'understanding': [456], '(e.g.,': [457, 1764], 'Proulx': [458], 'Servedio': [460], '2009;': [461, 466], 'Barton': [462, 623, 627, 635, 650, 804, 838, 2108, 2527, 2708, 2754], 'de': [464], 'Cara': [465], 'van': [467], 'Doorn': [468], '2009).': [471], 'A': [472, 493, 1413, 1683, 2022, 2047, 2215, 2316, 2361, 2365, 2439, 3076, 3275, 3802, 3825, 3858, 3939], 'consequence': [473], 'growth': [476, 2420], '"balkanization"': [478], 'subject': [481], '(Kirkpatrick': [482], 'Ravigne': [484], '2002),': [485], 'makes': [487, 543], 'synthetic': [488, 572], 'work': [491, 1173, 1487, 2618], 'important.': [492], 'major': [494], 'synthesis': [497], 'is,': [498, 592, 747, 762, 2299], 'however,': [499], 'modeling': [504], 'studies': [505, 547, 605], 'focus': [506, 566, 1474], 'on': [507, 567, 606, 945, 1055, 1188, 1254, 1632, 3551, 3572], 'conditions': [508, 526, 585], 'occur.': [512, 1550], 'Given': [513], 'inherent': [515], 'complexity': [516, 2498], 'speciation,': [518, 560, 590], 'usually': [521], 'difficult': [522, 548, 1161], 'extract': [524], 'analytical': [525, 800, 852, 1304, 1480, 2557, 3224, 3664, 4240, 4297], 'from': [529, 708, 748, 1070, 1913, 2007, 2044, 2514, 2722, 2915, 3325, 3971], 'realistic': [530], 'models.': [531, 867, 1426], 'Therefore,': [532], 'modelers': [533], 'often': [534], 'computer': [536, 3061], 'simulations': [537, 4238], 'limited': [539, 905], 'parameter': [540, 859, 4452], 'values.': [541], 'impossible.': [550], 'Thus,': [551, 1283, 2172, 3758], 'an': [554, 675, 754, 973, 2095, 3083], 'important': [555, 870], 'question': [556], 'what': [557, 3777], 'aspect': [558], 'if': [561, 773, 1599, 1622, 1835, 1888, 2545, 2871, 3415, 4013, 4488], 'not': [562, 882, 1118, 1286, 1549, 1555, 1674, 1813, 1822, 2613, 2759, 3549, 3701, 3794, 4157], '"occurrence,"': [563], 'get': [569], 'view.': [573], 'would': [575], 'like': [576], 'direct': [578], 'attention': [579], 'consequences': [581], 'of,': [582], 'rather': [583, 1327], 'than': [584, 727, 1923, 2540, 3087, 3149, 4443, 4457], 'for,': [586], 'isolation': [588, 602, 733, 931, 1012, 2376], 'strong': [594], 'reduction': [598, 1949], 'given': [601, 1067, 1877, 2944, 3046, 3485, 3615, 4133, 4226, 4379], 'mechanism.': [603], 'Previous': [604], 'found': [609, 1175, 1212], 'concept': [611, 819, 1042, 1214, 1972], 'rate"': [616], 'useful': [617, 1222, 1315], 'purpose': [620, 966], '(Bengtsson': [621, 802, 836, 2525], '1985;': [622, 803, 837, 2526], 'Bengtsson': [625, 806, 840, 899, 2029, 2110, 2529, 2553, 2710, 2992, 3494], '1986;': [626, 807], 'Gale': [629], '1993;': [630], '1997;': [632, 636, 1228, 1267], 'Piálek': [633], 'Cruzan': [639], '1998;': [640], '2002).': [644, 1269], 'According': [645, 2967, 4472], 'definition': [648, 1239, 1374, 2115], 'Bengtsson,': [652, 2756], 'me': [657, 707], 'results': [663, 735, 828, 3065, 3212, 3581, 3645, 4234, 4285], 'allele': [669, 1560, 1576, 1709, 1991, 1997, 2014, 2153, 2179, 2413, 2438, 2471, 2513, 2543, 2697, 2744, 3038, 3075, 3611, 3748, 3782, 3824, 4113, 4438], 'frequency': [670, 1561, 1707, 1721, 1989, 2436, 2544, 2740, 3039, 3612, 3831, 3969, 4005], 'change': [671], 'as': [672, 699, 1026, 1368, 1614, 1905, 1978, 2025, 2052, 2149, 3162, 3254, 3600, 3904, 4258], 'observed': [673], 'idealized': [676], 'reference': [677], 'model': [678, 2458, 2641, 2752, 2764, 2831, 4093, 4368], 'without': [679, 1278], 'any': [680, 696, 1517, 1677, 3703], 'kind': [681, 1678], 'class': [683], 'structure': [684, 1259, 3705, 4088], 'Then,': [688, 1380, 2163], 'modification': [691], 'be': [693, 1133, 1221, 1446, 1620, 1630, 1675, 2133, 2429, 2978, 3459, 3767, 3820, 4182], 'measured': [694], 'biological': [697], 'factor': [698], 'deviation': [701, 1907, 3965], 'actual': [710, 1884, 1915, 1925], 'm.': [713], 'Gene': [714], 'enhanced': [717], '(reduced)': [718], 'when': [719, 3115], 'higher': [725, 1922, 4441], '(lower)': [726], 'real': [729], 'rate.': [731, 1363, 1379, 1743, 1917, 2240, 3576, 3914], 'Reproductive': [732], 'typically': [734, 1947], 'reduction.': [739], 'calculation': [741], 'point': [751, 2648], 'view,': [753, 1001], 'eigenvalue': [755], '(Kobayashi': [757], '2008),': [760], 'unfortunately,': [763], 'only': [764, 883, 1119, 2331, 3873], 'tractable': [765], 'numerically': [766], 'most': [768, 1003], 'speciation.': [771, 1453, 2682], 'However,': [772, 1589, 1824, 2112, 3764], 'degree': [775, 1104, 1803, 1872, 4210], 'migrational': [777], 'mixture': [778], 'sufficiently': [780, 2549, 2876, 3423], 'system': [784, 1518, 1598, 2442, 2701, 3117, 4346], 'under': [785, 1208, 1676, 1897, 2256, 3196, 3629, 3646, 3687, 3716, 4273], 'consideration': [786], 'has': [787, 1131, 3118, 3568], 'certain': [789], 'level': [790], 'symmetry,': [792], 'systematic': [793], 'available': [796], 'obtain': [798, 826], 'good': [799], 'approximations': [801, 2558, 3225, 3665, 4298], 'Rousset': [808, 2067, 2073], '1999,': [809], '2008).': [814], 'founders': [816], 'made': [820], 'such': [823, 2432, 2598, 3180], 'analytic': [827], 'broad': [831], 'range': [832], '1986).': [841, 2030, 2111, 2530], 'Those': [842], 'great': [847], 'theory,': [850, 1318], 'because': [851, 922, 2333], 'formulae': [853, 1442, 2235, 2630, 2655], 'effect': [856, 1184, 1251, 3571], 'each': [858, 1605, 1612, 2301, 2591, 2596, 2748, 2837, 2856, 2910, 3301, 3341, 4068], 'separately': [860], 'allow': [863, 1624], 'It': [868, 1130, 3915], 'remark': [872, 1636, 2684, 3101], 'populations': [886, 938, 2118, 2704, 3407, 3694], 'but': [887, 1126, 3320, 3793], 'backgrounds': [892], 'context': [896, 918], '(Barton': [897, 2027], '1986),': [900], 'although': [901, 2686], 'applications': [902], 'been': [904], 'cases': [908, 3372], 'so': [909, 1528], 'far.': [910], 'generality': [912], 'respect': [914, 1773, 4374], 'particularly': [920, 1314, 1421, 2120], 'interesting,': [921], 'there': [923, 1536, 1654, 3296, 3303, 4324, 4430], 'now': [925, 2948, 4181], 'evidence': [929], 'may': [932, 1586, 1671, 1673, 1821, 2092], 'evolve': [933], 'even': [934], 'highly': [936], 'panmictic': [937], '(Via': [939], '2001).': [940], 'For': [941, 2075, 2451, 2766, 2884, 2919, 3451, 3776, 4245], 'critical': [943], 'view': [944], 'importance': [947], 'see': [951, 2459, 2778], 'Bolnick': [952], 'Fitzpatrick': [954, 956], '(2007),': [955], '(2008),': [959], 'Butlin': [961], '(2008).': [964, 2090], 'alternative': [974], 'approach': [975, 2536], 'quantifying': [977], 'take': [984], 'standpoint': [986], 'co-': [990], 'reciprocally': [992], 'adapted': [993], 'genes.': [996], 'Once': [997], 'admit': [999], 'natural': [1004, 1968], 'way': [1005], 'measure': [1015], 'shuffling': [1020], 'via': [1023, 2282], 'events,': [1025, 4009], 'main': [1030], 'source': [1031], 'breaking': [1033], 'up': [1034], '(Butlin': [1037], '2005).': [1038], 'introduce': [1040, 1372], 'gene-shuffling': [1053, 1805, 1874, 1951], 'based': [1054], 'at': [1058, 1544, 1604, 1690, 1699, 1711, 2003, 2036, 2181, 2209, 2385, 2422, 2808, 2864, 3281, 3292, 3300, 3750, 3786, 3882, 3933, 3952, 4018, 4077, 4153], 'coefficient': [1061, 1731, 2145, 2882, 4397], 'disequilibrium': [1064, 1193, 1734, 2148, 4465, 4477, 4506], 'loci': [1068, 1547, 1647, 1663, 1670, 1738, 1895, 3044, 3269, 3285], 'decreases': [1069], 'generation': [1072, 1712, 2276, 2838, 4057], 'next.': [1075], 'naturally': [1091], 'On': [1108, 2394], 'other': [1110, 1272, 1650, 1798, 2057, 2396, 2519, 2917, 3322], 'hand,': [1111, 2397], 'contains': [1116], 'information': [1117], 'about': [1120, 2771], 'amount': [1122, 1190], 'its': [1128, 1402, 1943, 3120], 'direction.': [1129], 'remarked': [1134], 'several': [1136, 2056], 'prior': [1137], 'authors': [1138, 1273], 'developed,': [1139], 'purposes,': [1142], 'similar': [1143, 3735], 'closely': [1145, 2121], 'related': [1146, 2122], 'concepts,': [1147], 'all': [1148, 2013, 2896, 3107, 3321, 3845, 4451], 'were': [1151, 3594], 'identically': [1152], 'called': [1153], 'rate."': [1157], 'Now,': [1158], 'attribute': [1163], 'origin': [1165], 'specific': [1169], 'study.': [1170], 'oldest': [1172], 'Golding': [1177], 'Strobeck': [1179], '(1980),': [1180], 'who': [1181], 'investigated': [1182, 4353], 'partial': [1186], 'selfing': [1187], 'generated': [1194], 'random': [1196, 1747, 1814, 1891, 2848, 2913], 'drift.': [1198], 'Nordborg,': [1199], 'B.': [1200], 'Charlesworth,': [1201], 'D.': [1203], 'Charlesworth': [1204, 1229], 'considered': [1205, 3740], 'coalescent': [1206], 'processes': [1207], 'balancing': [1209], '(Nordborg': [1223], '1996;': [1226], 'Nordborg': [1227], '1997).': [1232, 3510], 'Dye': [1233], 'Williams': [1235], '(1997)': [1236], 'adopted': [1237], 'analogous': [1245, 2096], 'ours': [1247, 2643], 'inbreeding': [1253], 'evolution': [1256], 'strain': [1258], 'parasites': [1261], '(see': [1262, 1952, 3507, 4161], 'Hastings': [1264], 'Wedgwood-Oppenheim': [1266], 'Lythgoe': [1268], 'Also,': [1270], 'loosely': [1274], 'used': [1275, 1447, 4250], 'term': [1277], 'strict': [1280], 'definition.': [1282], 'claiming': [1287], 'originality': [1289], 'concept.': [1292], 'Instead,': [1293, 3709], 'goals': [1295], 'provide': [1299, 2625], 'set': [1301, 2482], 'general': [1303, 1392, 2662, 3684, 3713, 3907], 'approximate': [1307], 'demonstrate,': [1322], 'applying': [1324, 1381], 'them': [1325], 'simple': [1328], 'examples,': [1329], 'analyzed': [1337], 'synthesized': [1339], 'within': [1340], 'framework.': [1344], 'As': [1345, 2736, 3470], 'byproduct,': [1347], 'reveal': [1349], 'relationship': [1353, 1458, 2231, 2637, 3897], 'structured': [1367], 'follows.': [1369], 'formula': [1394, 1416, 2249, 2664, 2675, 2998, 3535, 3547, 3686, 3715, 3760, 3909, 4243], 'demonstrate': [1401, 1439], '(Weak': [1410, 1434], 'Migration': [1411, 2242], 'Approximation).': [1412, 1436], 'derived': [1418, 2250, 2556, 3682], 'powerful': [1422], 'Again,': [1427], 'Recombination': [1435, 1504, 3671], 'compare': [1449], 'Finally,': [1454], 'discuss': [1456, 2635], 'studies,': [1464], 'possible': [1466, 2504], 'implications': [1467], 'data': [1469], 'analysis.': [1470], 'confine': [1472], 'presentation': [1477], 'approximation,': [1481], 'leave': [1483], 'future': [1486], 'apply': [1489], 'complex': [1496], 'involving': [1498], 'extensive': [1499, 3060], 'numerical': [1500, 3211, 3580, 3644, 4237, 4247, 4284, 4415], 'calculations.': [1501], 'Effective': [1503], 'Rate': [1505], 'BASIC': [1506], 'ASSUMPTIONS': [1507], 'developed': [1510], 'section': [1513, 2253], 'applicable': [1515, 1570, 1593, 2255], 'satisfying': [1519], 'following': [1521, 2137, 2258, 2661, 2834], 'conditions:': [1522], 'population': [1525, 1762, 2261, 2419, 2721, 2725, 2749, 3398], 'size': [1526], 'large': [1529], 'drift': [1532], 'negligible,': [1534], 'diallelic': [1539, 2129, 3268], 'loci,': [1540, 1651, 1878], 'mutation': [1543, 1633], 'does': [1548, 3548, 4156], 'If': [1551, 1744, 1810, 4499, 4519], 'condition': [1552, 2929, 3182, 4500, 4520], 'met': [1556], 'demography': [1558], 'dynamics': [1562, 1573, 3040, 3613], 'include': [1563], 'stochastic': [1564], 'factors,': [1565], 'expected': [1575], 'frequencies.': [1577], 'assumption': [1579, 2338, 3689, 3718, 3732, 4172], 'alleles': [1583, 1603, 1689, 2315, 3280, 3291, 3801, 4329], 'per': [1584], 'locus': [1585, 1606, 1693, 2006, 2039, 2195, 2212, 2388, 2400, 2408, 2411, 2811, 2867, 2957, 3445, 3455, 3789, 3885, 3936, 3955, 4021, 4049, 4143, 4155, 4319, 4327, 4337, 4484], 'seem': [1587], 'restrictive.': [1588], 'multiallele': [1597], 'divide': [1601], 'groups': [1609, 2360], 'regard': [1611], 'group': [1613, 2009, 2018, 2046, 2217, 3860], '"allele."': [1616], 'Condition': [1617], 'dropped': [1621], 'dependent': [1631], 'rates.': [1634, 2903], 'none': [1638], 'assumptions': [1641, 3250], 'restricts': [1642], 'focal': [1646, 1720], 'interact': [1648], 'no': [1656, 1749, 1898], 'limitation': [1657], 'total': [1660, 3306, 3838], 'number': [1661, 3839, 3853], 'system.': [1666, 2735], 'Further,': [1667, 3800], 'selection.': [1680, 1899, 2393, 4335, 4403], 'DEFINITION': [1681], 'Let': [1682, 1703, 1725, 1985, 2284, 3274, 3816, 4043], 'denote': [1686, 1705, 1729, 1861, 1987, 2288, 3278], 'B': [1695, 2414], 'b': [1697], 'those': [1698], 'locus.': [1702, 2802, 2966, 3450, 3469, 3730, 4471], 'ϕx(t)': [1704], 'x': [1710, 1992], 't.': [1713], 'omit': [1715], 'argument': [1716, 2568], 't': [1717], 'whenever': [1718], 'temporally': [1723, 2161, 2177], 'constant.': [1724, 2162], 'D(t)': [1726, 1753, 1839], '=ϕAB(t)': [1727], '−ϕA(t)ϕB(t)': [1728], 'r': [1740, 1775, 1800, 1887, 3724], 'acting,': [1752, 1818], 'follows': [1754], 'classical': [1759, 4092], 'result': [1760, 2986, 3489, 4461], 'genetics': [1763], 'Crow': [1765, 3958], 'Kimura': [1767, 3960], '1970).': [1768], 'Solving': [1769], 'equation': [1770, 1819, 1834, 2165, 2168, 2447, 2491, 2657, 3141, 3228, 3668, 3902, 4035, 4187, 4190, 4301], 'yields': [1776, 2170, 2659, 3005, 3542, 4036], 'Equation': [1778, 3007, 4196], 'shows': [1780, 3009, 3092, 3223, 3579, 3663, 4198, 4232, 4296], 'equal': [1786, 1881, 2197, 2350, 2899], 'per-generation': [1790], 'decreasing': [1791, 3017, 4206], 'disequilibrium.': [1796], 'In': [1797, 2223, 2531, 2621, 2785, 2829, 2859, 3676, 3697, 4011, 4064, 4110], 'words,': [1799], 'loci.': [1809, 2130, 3247, 4498], 'hold.': [1823], 'calculate': [1827, 2506, 3437, 3514], 'right-hand': [1829], 'side': [1830], 'above': [1833, 2114, 2464, 3095, 3546, 4034], 'values': [1837, 3527, 3593], 'D(t+': [1841], '1)': [1842, 1930, 1941], 'known.': [1844], 'call': [1846, 2270, 2405], 'quantity': [1848], 'A/a': [1855, 2183, 2191, 2387, 2810, 2866, 2956, 3444, 3465, 3727, 3788, 3888, 3954, 4020, 4048, 4142, 4318, 4468, 4480], 'B/b': [1858, 2005, 2194, 2211, 2399, 2407, 2965, 3449, 3454, 3729, 3884, 3935], 'locus,': [1859, 2184, 3302], 're(t)': [1864], 'define': [1901, 2094], '"gene': [1902], 'shuffle': [1903], 'modification"': [1904], '(re(t)/r': [1928, 1939], '>': [1929, 3988, 4393], 'indicates': [1931, 1942, 3979], 'enhancement': [1932], 'gene-shuffling,': [1934], 'whereas': [1935, 2815, 3982], 'opposite': [1937, 2042], 'situation': [1938, 3238, 3736], '<': [1940, 3984], 'suppression.': [1944], 'involves': [1946], 'discussed': [1954, 3109], 'below).': [1955], 'RELATIONSHIP': [1956], 'TO': [1957], 'MIGRATION': [1960], 'extension': [1969], 'will': [1980], 'show': [1981], 'section.': [1984, 3708], 'ϕx|y(t)': [1986], 'among': [1993], 'gametes': [1995, 2011, 2020, 3855], 'having': [1996, 2012, 2021, 4439], 'y.': [1998], 'defined': [2024, 2051, 3139], 'Likewise,': [2031], 'direction': [2043], 'gamete': [2045, 2216, 2221, 2359, 2476, 3814, 3833, 3859, 3889, 3941], '(5)': [2053], 'There': [2054, 4421], 'ways': [2058], 'defining': [2060], '(Petry': [2065], '1983;': [2066], '1999;': [2068], '2002;': [2072], 'comparison': [2077, 3426], 'definitions,': [2081], 'refer': [2083], 'reader': [2085], 'describe': [2099], 'across': [2102], 'spatial': [2104], 'continuum': [2105], '(Nagylaki': [2106], '1976;': [2107], 'seen': [2134], 'reasoning.': [2138], 'Note': [2139, 3359, 3850, 3997], 'rewrite': [2143, 3901], '(6)': [2150, 2166], 'Suppose': [2151, 2313, 4054], 'frequencies': [2154, 2180, 3073, 3785, 3822, 4114, 4257, 4513], 'ϕA(t)': [2155], '=ϕA': [2156], 'ϕa(t)': [2158], '=ϕa': [2159], 'substituting': [2164, 4186], '(7)': [2171, 2448, 2658], 'case': [2175, 2761, 2922, 3499, 3585, 4364], 'constant': [2178, 2444, 2746, 4119], 'sum': [2200], 'rates': [2205, 2291, 2344, 2355, 2535, 2547, 2874, 2891, 3138, 3194, 3418, 3627, 4271], 'group.': [2222, 3942], 'next': [2225], 'sections,': [2227], 'Weak': [2241, 3670], 'Approximation': [2243, 3672], 'GENERAL': [2244, 3673], 'FORMULA': [2245, 3674], 'I': [2246], 'assumptions.': [2259], 'consists': [2262], 'subpopulations': [2265], 'parapatry,': [2267], '"demes"': [2271], '1': [2272, 2294, 2311, 2324, 2584, 2827, 3399, 3483, 3978, 3985, 3989], '2.': [2274, 3408, 3524], 'Every': [2275], 'demes': [2279, 2323, 2348, 2425, 2474], 'exchange': [2280], 'individuals': [2281, 2806, 2817, 3841, 3878, 4017, 4072], 'migration.': [2283], 'm': [2285, 2307, 3419], 'm*': [2287, 3421], 'immigration': [2290, 2873, 2902], 'deme': [2293, 2310, 2516, 2583, 2597, 2857, 2911, 3482, 3523], '2,': [2296, 2326], 'respectively.': [2297, 3219, 3652, 3996, 4010, 4292], 'That': [2298], 'after': [2300], 'event,': [2303], 'immigrants': [2304, 2581, 2908], 'constitute': [2305], 'fraction': [2306], '(m*)': [2308], '(2).': [2312, 2585], 'nearly': [2320], 'fixed': [2321], 'respectively': [2327], '(fixation': [2328], 'attained': [2330], 'nearly,': [2332], 'continuous': [2335], 'migration).': [2336], 'ensures': [2339], 'a.': [2363], 'Alleles': [2364], 'comprise': [2368], 'part': [2370], 'causing': [2374, 4333], 'demes,': [2380], 'differentiation': [2384, 2863, 2934, 3411], 'maintained': [2390, 2870, 3414, 4487], 'selectively': [2402, 2960, 4145], 'neutral.': [2403], 'marker': [2410, 2416, 2470, 2512, 2542, 2696, 2743, 3089, 3747], 'allele.': [2417, 3090], 'supposed': [2427, 3457, 3806], 'same,': [2431], 'overall': [2435], 'stays': [2443, 2745], 'hence': [2446], 'applicable.': [2450, 3099], 'rigorous': [2453, 2627], 'description': [2454, 3031], 'underlying': [2457], 'Appendix': [2460, 2622, 2779, 2969, 3048, 3473, 3617, 4228], 'S1.': [2461, 2780], 'Under': [2462, 4170], 'assumptions,': [2465], '(or': [2475], 'groups)': [2477], 'described': [2479, 3335], 'difference': [2484, 3133], 'equations': [2485, 3035, 3126, 3607, 4224], '(Appendix': [2486, 2551], 'S1)': [2487], 'matrix': [2490, 3338, 3345, 3361], '(Appendices': [2492], 'S1': [2493], 'S2).': [2495, 2552], 'Despite': [2496], 'dynamics,': [2501], 'general,': [2532], 'equilibria': [2537], 'much': [2538, 3085], 'faster': [2539, 3086], 'small': [2550, 2877, 4174], '(1985)': [2554, 2640, 3495], 'rates,': [2562], 'showed': [2564], 'heuristic': [2567], '(8)': [2570, 2631], '(9)': [2571], 'where': [2572, 3340, 3745, 3919, 4369, 4382, 4490], 'v': [2573, 3529, 3923], '(v*)': [2574, 3924], 'average': [2577, 2601, 2972, 3476, 3516, 3927], 'value': [2579, 2589, 2603, 2974, 3130, 3156, 3177, 3478, 3518, 3929, 4125], 'Here,': [2586, 2646, 3922, 4359], 'individual': [2592, 4069], 'normalized': [2594], 'residents': [2605], '(note': [2608], 'normalization': [2611], 'was': [2612, 2935, 2988, 3490, 4097, 4347, 4407], 'taken': [2614], 'original': [2617], 'Bengtsson).': [2620], 'S2,': [2623], 'proof': [2628], '(9),': [2633], "Bengtsson's": [2639], 'detail.': [2645], 'out': [2649, 2718, 2730], 'substitution': [2651], 'rate:': [2669], '(10)': [2670, 3004, 3541, 3761, 3903], 'Below,': [2671], 'analyze': [2677, 2790], 'consider': [2688, 3236, 3386, 4361], 'closed': [2700], 'connected': [2705], 'migration,': [2707, 2845], '(1986)': [2711], 'assumed': [2712, 3170, 4139], 'influx': [2715], '(efflux': [2717], 'of)': [2719], '(into)': [2723], 'another': [2724], 'balanced': [2727], 'efflux': [2729], '(influx': [2732], 'into)': [2733], 'result,': [2738], 'setting.': [2765], 'detailed': [2769], 'discussion': [2770], 'differences': [2773], 'EXAMPLE': [2781, 3230, 4039, 4303], '1:': [2782], 'SINGLE-LOCUS': [2783], 'INCOMPATIBILITIES': [2784], 'example,': [2788], 'scenario': [2792, 3388], 'incompatibilities': [2796, 3242, 3263, 3368], 'caused': [2798, 3244, 3265], 'assume': [2804, 2887, 2906, 3260, 3311, 3702, 3780, 4045, 4312], 'homozygous': [2807, 4014], 'fitness': [2813, 2826, 3318, 3348, 4371, 4405, 4427, 4442, 4456], 'one,': [2814], 'heterozygous': [2816, 3870, 3877, 4016], 'genotype': [2819, 3351, 3391, 3401, 3784, 3968, 4076, 4152, 4389], 'Aa': [2820], 'reduced': [2823, 3327], 'viability,': [2824], '−s.': [2828], 'setting,': [2832], 'events': [2835, 3875], 'occur': [2836, 4061], 'order:': [2840], 'viability': [2842, 3328, 3561, 4339, 4387], 'selection,': [2843], 'reproduction.': [2851], 'latter': [2853], 'occurs': [2854], 'separately.': [2858], 'setup,': [2861], 'stably': [2869, 3413, 4486, 4508], 'compared': [2878], 's.': [2883], 'simplicity,': [2885, 3452], 'emigration': [2890], 'genotypes': [2897, 3299, 3313, 3323], 'further': [2905], 'samples': [2914], 'deme.': [2918], 'special': [2921, 3371, 3498, 3584, 4363], 'symmetric': [2924, 3171, 3200, 3633], '(m=m*),': [2926, 3173], 'exact': [2928], 'stability': [2931], 'obtained': [2936, 2990, 3492], 'Karlin': [2938], 'McGregor': [2940], '(1972)': [2941], '(11)': [2946, 3183], 'determine': [2949, 3114], 'neutral': [2961, 3448, 4146], 'physically': [2963, 3272, 3460], 'unlinked': [2964, 3461], 'S3,': [2970, 3474], 'migrants': [2976, 3480, 3521], 'calculated': [2979, 3226, 3666, 4183, 4299], 'analytically': [2980], 'computes': [2982, 4193], '(12)': [2984], '(1985).': [2993], 'Substitution': [2994], '(13)': [3006, 3008], 'function': [3018, 4207, 4372, 4406], 's': [3020, 3179, 4392], 'converges': [3022], 'zero': [3024], 's→': [3026], '1.': [3027, 3070, 4029, 4220], 'precise': [3029], 'recursion': [3034, 3125, 3606, 4223], 'S4.': [3049, 3618, 4229], 'To': [3050, 3113], 'test': [3051], 'accuracy': [3053], 'formula,': [3057], 'performed': [3059], 'simulations.': [3062], 'Some': [3063], 'characteristic': [3064, 4233], 'shown': [3067, 3471], 'Figure': [3069, 3186, 3577, 3619, 4230, 4263], 'Although': [3071], 'vary': [3077], 'time,': [3079, 4121], 'they': [3080], 'converge': [3081], 'equilibrium': [3084, 3164, 4124, 4256, 4512], 'technique': [3097], 'true': [3105], 'article.': [3112], 'reached': [3119], 'equilibrium,': [3121, 3792], 'iterated': [3123], 'until': [3127], 'absolute': [3129], 'successive': [3145], 'generations': [3146], 'becomes': [3147], 'lower': [3148, 4455], '10−7,': [3150], 'then': [3152], 'took': [3153], 'final': [3155], 'value.': [3165], 'Throughout': [3166], 'simulations,': [3168], 'chose': [3175], 'fulfilled.': [3185], '1Open': [3187], 'figure': [3189, 3622, 4266], 'viewerPowerPoint': [3190, 3623, 4267], 'Normalized': [3191, 3624, 4268], '(re/(m+m*))': [3195, 3628], '(m=m*).': [3202, 3635], 'Filled': [3203, 3636, 4276], 'circles,': [3204, 3206, 3637, 3639, 4277, 4279], 'open': [3205, 3208, 3638, 3641, 4278, 4281], 'squares': [3209, 3642, 4282], 'indicate': [3210, 3643, 3990, 4283], 'm=': [3214, 3647, 3894], '0.001,': [3215, 3648], '0.01,': [3216, 3649, 4288], '0.05,': [3218, 3651], 'solid': [3221, 3661, 4294], 'line': [3222, 3662, 4295], '(13).': [3229], '2:': [3231], 'TWO-LOCUS': [3232], 'INCOMPATIBILITY': [3233], 'Next,': [3234, 4310], 'basic': [3249], 'example.': [3258, 3604], 'unlinked.': [3273], 'E': [3287, 4330], 'e': [3289, 4332], 'other.': [3294], 'Because': [3295, 3872], 'three': [3298], 'nine': [3308], 'genotypes.': [3309], 'AAEE': [3314, 3392], 'aaee': [3316, 3402], 'suffer': [3324], 'due': [3329], 'incompatibility.': [3332], 'mathematically': [3336], '(14)': [3339], 'element': [3342], 'represents': [3346], 'designated': [3352], 'row': [3355], 'column': [3357], 'labels.': [3358], 'H': [3362], 'reduces': [3363], 'well-known': [3366], 'Dobzhansky–Muller': [3367, 3501], 's3=s6=s7=': [3374], '0': [3375, 3378, 3506, 4217, 4394], 's1=s2=s5=': [3377], '(Dobzhansky': [3379], '1937;': [3380], 'Muller': [3381], '1942;': [3382], 'close': [3394, 3403], 'fixation': [3396, 3405], 'low': [3424, 3721, 3752], 'coefficients': [3430], 's1,...,': [3431], 's7.': [3432], 'Our': [3433], 'aim': [3434], 'E/e': [3468, 4470, 4483], '(15)': [3487], 'incompatibilities,': [3502], 's3=': [3505], 'likewise': [3513], 'v*': [3519, 3531], 'Substituting': [3525, 4030], '(16)': [3543], 'Interestingly,': [3544], 'depend': [3550], 's2': [3552], 's6.': [3554], 'Another': [3555], 'interesting': [3556], 'observation': [3557], 'first-generation': [3564], 'AaEe': [3566], '(s4)': [3567], 'dominant': [3570], '2': [3578], 's=s2=s6=s4,': [3587], 'σ=': [3588], '0.5s=s1=s3=s5=s7,': [3589], 'm=m*.': [3591], 'Equilibrium': [3592], 'determined': [3595], 'using': [3596, 4242], 'criterion': [3599], 'Complete': [3605], 'describing': [3608], 'entire': [3610], '2Open': [3620], 'Other': [3653], 'parameters': [3654], 's2=s4=s6=s': [3656], 's1=s3=s5=s7=': [3658], '0.5s.': [3659], '(16).': [3669], 'II': [3675], 'section,': [3679, 3744], 'low.': [3696], 'contrast,': [3698], 'do': [3700], 'creates': [3733], 'migrates': [3749], 'backgrounds.': [3757], 'holds.': [3763], 'must': [3766], 'rewritten': [3768], 'terms': [3770], 'r.': [3775], 'follows,': [3778], 'necessarily': [3795], 'Hardy–Weinberg': [3798, 3973], 'equilibrium.': [3799, 3974], 'cause': [3808], 'isolating': [3810], 'groups.': [3815], 'ϕ+A': [3817], 'ϕ+a': [3819], 'a,': [3827], 'respectively,': [3828], 'ϕ+Aa': [3829], 'Aa,': [3834], 'n+': [3836], 'population,': [3844], 'assessed': [3846], 'just': [3847], 'before': [3848, 3861, 4342], '2n+ϕ+A,': [3864], 'n+ϕ+Aa': [3867], 'individuals.': [3871], 'contribute': [3879], 'groups,': [3890], 'holds': [3892, 3916, 4026, 4167], '(ϕ+Aa/2ϕ+A)r.': [3895], '(17)': [3918], '(18)': [3920], '(19)': [3921], '"migrant"': [3931], '(a)': [3940], 'symbol': [3944], 'π': [3945, 3983, 3987, 4000, 4131], 'stands': [3946], '"index': [3949], 'panmixia"': [3951], '(Wright': [3956], '1951;': [3957], '1970),': [3961], 'distribution': [3970], 'Roughly': [3975], 'speaking,': [3976], 'π=': [3977], 'panmixia': [3981, 4130], 'assortment': [3991], 'disassortment': [3993], 'alleles,': [3995], 'account': [4001], 'impact': [4003], '"effective"': [4007], 'particular,': [4012], 'equally': [4023], 'fit,': [4024], 'v=v*=': [4028], '(20)': [4038], '3:': [4040], 'ASSORTATIVE': [4041, 4305], 'MATING': [4042, 4306], 'codes': [4050], 'mating.': [4053, 4275], 'every': [4056], 'reproduction': [4060, 4066], 'order.': [4063], 'phase,': [4067], 'mates': [4070, 4084], 'A/a-locus': [4079], 'probability': [4081], 'α,': [4082], 'randomly': [4085], 'otherwise.': [4086], 'Geographic': [4087], 'absent.': [4090], 'introduced': [4099, 4348, 4408], 'Wright': [4101], '(1921)': [4102], 'studied': [4104], 'detail': [4106, 4355], "O'Donald": [4108, 4138], '(1960).': [4109], "O'Donald's": [4111], 'model,': [4112], 'ϕA': [4115], 'ϕa': [4117], '(21)': [4135, 4188], "(O'Donald": [4136], '1960).': [4137], 'sense': [4149], 'affect': [4158], 'success': [4160], 'Hence,': [4165], '.': [4169], 'r,': [4175], 'easily': [4184], '(20),': [4191], '(22)': [4195, 4197], 'α': [4214], 're=': [4216], 'α=': [4219], 'complete': [4222], 'solutions': [4241], '(22).': [4244, 4302], 'calculations,': [4248], 'criteria': [4253], 'determining': [4255], 'examples.': [4262], '3Open': [4264], '(re/r)': [4272], 'r=': [4287], '0.1,': [4289], '0.5,': [4291], '4:': [4304], 'WITH': [4307], 'DISRUPTIVE': [4308], 'SELECTION': [4309], 'that,': [4313], 'addition': [4315], 'coding': [4320], 'mating,': [4323], 'affects': [4338], 'acts': [4341], 'Udovic': [4350, 4367, 4419], '(1980)': [4351], '(2003).': [4358], '(23)': [4381], 'wx': [4383], 'x,': [4390], '(2004)': [4411], 'perform': [4413], 'analysis': [4416], 'model.': [4420], 'remarks': [4424], 'regarding': [4425], 'function.': [4428], 'First,': [4429], 'minority': [4433], 'rare': [4437], 'common': [4445], 'type.': [4446], 'Second,': [4447], 'heterozygotes': [4448], 'constellations': [4453], 'homozygotes.': [4458], "Gavrilets'": [4474], 'analysis,': [4475], '(24)': [4489, 4501, 4521], 'fulfilled,': [4503], 'maintained,': [4509], 'ϕA=ϕa=ϕE=ϕe=': [4515], '1/2': [4516], '(Udovic': [4517], '1980).': [4518]}, 'cited_by_api_url': 'https://api.openalex.org/works?filter=cites:W1553570292', 'counts_by_year': [{'year': 2022, 'cited_by_count': 1}, {'year': 2019, 'cited_by_count': 1}, {'year': 2016, 'cited_by_count': 1}, {'year': 2013, 'cited_by_count': 2}, {'year': 2012, 'cited_by_count': 1}], 'updated_date': '2024-12-14T11:04:58.069791', 'created_date': '2016-06-24'}